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Serum levels of VEGF and HIF-1α are significantly elevated in PsA patients compared to healthy controls or PsA patients in remission (187) suggesting a direct effect of the HIF-1α factor in the increased expression of VEGF which induces proliferation, migration, and differentiation of endothelial cells (188, 189)

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Furthermore, at sub-cytotoxic concentrations, the size of the colonies was reduced, suggesting an effect on cell proliferation